microscopio cabeceras

Luis María  Carrascal de la Puente


915668850 - 988963




Profesor de investigación del CSIC

Research lines



Luis M. Carrascal is a Research Professor at the Department of Biogeography & Global Change of the Museo Nacional de Ciencias Naturales (Spain). He is also a member of the Integrative Ecological Biogeography research group. 

His current research interests are focused on macroecology (mainly related to biogeographical patterns of the avifauna of the South-Western Palearctic), and on the analysis of the interaction between abundance-distribution patterns of birds and some autoecological traits such as body condition, foraging ecology and responses to predation risk.

His research program includes two specific scientific goals.


  • The first is to explore the determinants of the geographical range and population density of terrestrial animal species at different spatial scales, accounting for ecological components while controlling for phylogenetic effects, and considering the relationships among the interspecific variation in population trends, distribution patterns, habitat preferences, niche breadth and morphology.
  • The second goal is to study the interaction between environmental food availability and the thermal state of the environment (also considering the climate variation in the recent years) as determinants of the body condition, foraging ecology, distribution, abundance of vertebrates and beetles inhabiting the Iberian peninsula.


He has published more than 80 scientific papers included in the ISI Web of Knowledge, has supervised nine PhD Thesis, and has been deeply involved in teaching statistics and scientific method for postgraduate students and technicians related to environmental research.



Links / Enlaces:

ResearcherID: B-8381-2008

ORCID: 0000-0003-1288-5531



Google Scholar

Home page




MAIN SCIENTIFIC PUBLICATIONS (since 1990) / Publicaciones científicas seleccionadas (desde 1990)


Carrascal, L.M.; Galván, I.; Sánchez-Oliver, J.S.; Rey-Benayas, J.M. 2014. Regional distribution predicts bird occurrence in Mediterranean cropland afforestations. Ecological Research 29: 203-211.


Villén-Pérez, S.; Carrascal, L.M.; Seoane, J. 2013. Foraging patch selection in winter: a balance between predation risk and thermoregulation costs. PLoS One 8(7): e68448. doi:10.1371/journal.pone.0068448.


Hortal, J.; Carrascal, L.M.; Triantis, K.A.; Thébault, E.; Meiri, S.; Sfenthourakis, S. 2013. Species richness can decrease with altitude, but not with habitat diversity. PNAS 110(24): E2149-50.


Carrascal, L.M.; Cayuela, L.; Palomino, D.; Seoane, J. 2012. What species-specific traits make a bird a better surrogate of native species richness? A test with insular avifauna. Biological Conservation 152:204-211.


Carrascal, L.M.; Santos, T.; Tellería, J.L. 2012. Does day length affect winter bird distribution? Testing the role of an elusive variable. PLOS One 7(2): e32733. doi:10.1371/journal.pone.0032733.


Seoane, J.; Carrascal, L.M.; Palomino, D. 2011. Assessing the ecological basis of conservation priority lists for bird species in an island scenario. J. Nature Conservation 19:103-115.


Rey Benayas, J.M.; Galván, I.; Carrascal, L.M. 2010. Differential effects of vegetation restoration in Mediterranean abandoned cropland by secondary succession and pine plantations on bird assemblages. Forest Ecology & Management 260:87-95.


Carrascal, L.M.; Galván, I.; Gordo, O. 2009. Partial least squares regression as an alternative to most currently used regression methods in Ecology. Oikos, 118:681-690.


Carrascal, L.M.; Seoane, J.; Palomino, D.; Polo, V. 2008. Explanations for bird species range size: ecological correlates and phylogenetic effects in the Canary Islands. Journal of Biogeography 35:2061–2073.


Seoane, J.; Carrascal, L.M. 2008. Interspecific differences in population trends of Spanish birds are related to habitat and climatic preferences. Global Ecology & Biogeography 17:111-121.


Polo, V.; Carrascal, L.M.; Metcalfe, N.B. 2007. The effects of latitude and day length on fattening strategies of wintering coal tits (Periparus ater L): a field study and aviary experiment. Journal of Animal Ecology 76:866-872.


Palomino, D.; Carrascal, L.M. 2007. Threshold distances to nearby cities and roads influence the bird community of a mosaic landscape. Biological Conservation 140:100-109.


Bosch, J.; Carrascal, L.M.; Durán, L.; Walker, S.; Fisher, M.C. 2007. Climate change and outbreaks of amphibian chytridiomycosis in a montane area of Central Spain; is there a link? Proceedings Royal Society London B 274:253–260.


Palomino, D.; Carrascal, L.M. 2006. Urban influence on birds at a regional escale. A case study with the avifauna of northern Madrid province. Landscape & Urban Planning 77:276-290.


Carrascal, L.M.; Polo, V. 2006. Effects of wing area reduction on winter body mass and foraging behaviour in coal tits: field and aviary experiments. Animal Behaviour 72: 663-672.


De La Montaña, E.; Rey Benayas, J.M.; Carrascal, L.M. 2006. Response of bird communities to silvicultural thinning of Mediterranean maquis. Journal of Applied Ecology 43:651–659.


Carrascal, L.M.; Alonso, C.L. 2006. Habitat use under latent predation risk. A case study with wintering forest birds. Oikos 112: 51-62.


Carrascal, L.M.; Díaz, L. 2006. Winter bird distribution in abiotic and habitat structural gradients. A case study with mediterranean montane oakwoods. EcoScience 13:100-110.


Seoane, J.; Carrascal, L.M.; Alonso, C.L. y Palomino, D. 2005. Species-specific traits associated to prediction errors in bird habitat suitability modelling. Ecological Modelling 185:299-308.


Carrascal, L. M.; Lobo, J. L. 2003. Respuestas a viejas preguntas con nuevos datos: estudio de los patrones de distribución de la avifauna española y consecuencias para su conservación. Pp. 645-662 y 718-721 en Martí, R., Del Moral, J.C. (Eds.). Atlas de las Aves Reproductoras de España. Dirección General de la Conservación de la Naturaleza-Sociedad Española de Ornitología, Madrid.


Belliure, J.; Carrascal. L.M. 2002. Influence of heat transmission mode on heating rates and on the selection of patches for heating in a Mediterranean lizard. Phys. & Bioch. Zool. 75(4):369-376.


Carrascal, L.M.; Díaz, J.A.; Huertas, D.L.; Mozetich, I. 2001. Behavioral thermoregulation by treecreepers: trade-off between saving energy and reducing crypsis. Ecology 82:1642-1654.


Carrascal, L.M.; Polo, V. 1999. Coal tits, Parus ater, lose weight in response to chases by predators. Animal Behaviour 58:281-285.


Polo, V.; Carrascal, L.M. 1999. Shaping the body size distribution of passeriformes: habitat use and body size are evolutionarily and ecologically related. Journal of Animal Ecology. 68:324-337.


Carrascal, L.M.; Senar, J.C.; Mozetich, I.; Uribe, F.; Domenech, J. 1998. Interaction between environmental stress, body condition, nutritional status and dominance in mediterranean great tits (Parus major) during winter. Auk 115:727-738.


Moreno, E.; Barbosa, A.; Carrascal, L.M. 1997. Should congruence between intra- and interspecific ecomorphological relationships be expected? A case study with the Great Tit Parus major. Proceedings of The Royal Society London: B 264:533-539.


Belliure, J.; Carrascal, L.M.; Díaz, J.A. 1996. Covariation of thermal biology and foraging mode in two mediterranean lacertid lizards. Ecology 77:1163-1173.


Carrascal, L.M.; Moreno, E.; Mozetich, I. 1995. Ecological plasticity of morphological designs. An experimental analysis with tit species. Canadian J. Zoology 73:2005-2009.


Carrascal, L.M.; Moreno, E.; Valido, A. 1994. Morphological evolution and changes in foraging behaviour of island and mainland populations of Blue Tit, Parus caeruleus. A test of convergence and ecomorphological hypoteses. Evolutionary Ecology 7:25-35.


Tellería, J.L.; Carrascal, L.M. 1994. Weight-density relationships between and within bird communities. Implications of niche space and vegetation structure. American Naturalist 141:1083-1092.


Moreno, E.; Carrascal, L.M. 1993. Leg morphology and feeding postures in four Parus species: an experimental ecomorphological approach. Ecology 74:2037-2044.


Carrascal, L.M.; Bautista, L.M.; Lázaro, E. 1993. Geographical variation in the density of the White Stork Ciconia ciconia in Spain: influence of habitat structure and climate. Biological Conservation 65:83-87.


Díaz, J.A.; Carrascal, L.M. 1993. Variation in the effect of profitability on prey size selection by the lacertid lizard Psammodromus algirus. Oecologia (Berl.) 94:23-29.


Carrascal, L.M.; López, P.; Martín, J.; Salvador, A. 1992. Basking and antipredator behaviour in a high altitude lizard: implications of heat-exchange rate. Ethology 92:143-154.


Carrascal, L.M.; Moreno, E. 1992. Proximal costs and benefits of heterospecific social foraging in Great Tit Parus major. Canadian Journal of Zoology 70:1947-1952.


Carrascal, L.M.; Tellería, J.L.; Valido, A. 1992. Habitat distribution of canary chaffinches among islands: competitive exclusion or species-specific habitat preferences? Journal of Biogeography 19:383-390.


Carrascal, L.M.; Tellería, J.L. 1991. Bird size and density: a regional approach. American Naturalist 138:777-784.


Díaz, J.A.; Carrascal, L.M. 1991. Regional distribution of a Mediterranean lizard: influence of habitat cues and prey abundance. Journal of Biogeography 18:291-297.


Carrascal, L.M.; Moreno, E.; Tellería, J.L. 1990. Ecomorphological relationships in a group of insectivorous birds of temperate forests in winter. Holarctic Ecology (Ecography) 13:105-111.



Ministerio de Ciencia, Innovación y UniversidadesConsejo Superior de Investigaciones Científicas

Web realizada con la
colaboración de